हस्ताज्जातौ

Operational rule vidhi

The affix इनि (इन्) comes in the sense of मतुप् after the word हस्त when a genus is denoted by the word so formed.

Word by word (padaccheda)

Left context हस्तात् pañcamī · ekavacanaRight context जातौ saptamī · ekavacana

Left context Pañcamī (5th case): "after X", the operation applies to what follows (1.1.67 तस्मादित्युत्तरस्य). Also used for limits and sources.
Right context Saptamī (7th case): "when X follows", the operation applies to what precedes (1.1.66 तस्मिन्निति निर्दिष्टे पूर्वस्य). Also a condition or domain ("in the sense of").

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Full reading

हस्तात् जातौ

Pāṇini leaves out words that carry over from earlier sūtras. Expand to see the rule as a reader actually understands it.

Commentary

The affix इनि (इन्) comes in the sense of मतुप् after the word हस्त when a genus is denoted by the word so formed.
Thus हस्तोऽस्यास्ति = हस्तिन् 'elephant'. Why do we say when meaning a जाति 'a genus' ? Observe हस्तवान् पुरुषः ॥

Śrīśa Chandra Vasu, The Ashtadhyayi of Panini (1897), via ashtadhyayi.com.