निष्कुलान्निष्कोषणे

Operational rule vidhi

The affix डाच् comes after the word निष्कुल followed by the verb कृ 'to make' when the sense is 'to disembowel'.

Word by word (padaccheda)

Left context निष्कुलात् pañcamī · ekavacanaRight context निष्कोषणे saptamī · ekavacana

Left context Pañcamī (5th case): "after X", the operation applies to what follows (1.1.67 तस्मादित्युत्तरस्य). Also used for limits and sources.
Right context Saptamī (7th case): "when X follows", the operation applies to what precedes (1.1.66 तस्मिन्निति निर्दिष्टे पूर्वस्य). Also a condition or domain ("in the sense of").

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Full reading

निष्कुलात् निष्कोषणे

Pāṇini leaves out words that carry over from earlier sūtras. Expand to see the rule as a reader actually understands it.

Commentary

The affix डाच् comes after the word निष्कुल followed by the verb कृ 'to make' when the sense is 'to disembowel'.
The verb कृ should be read into the sutra. The word निष्कोषण means to bring the inner organs out, or to eviscerate. Thus निष्कुल करोति पशून् "he disembowels or guts the animals". निष्कुला करोति दाडिमम् "he takes out the kernel or seeds of the pomegranite". Why do we say "when meaning to disembowel or take out the kernel"? Observe निष्कुलान् करोति शत्रून् "he makes the enemies family-less".

Śrīśa Chandra Vasu, The Ashtadhyayi of Panini (1897), via ashtadhyayi.com.