4.1.55 नासिकोदरौष्ठजङ्घादन्तकर्णशृङ्गाच्च

Word by word (padaccheda)

Left context नासिक- उदर- ओष्ठ- जङ्घा- दन्त- कर्ण- शृङ्गात् pañcamī · ekavacanaIndeclinable च avyaya

Left context Pañcamī (5th case): "after X", the operation applies to what follows (1.1.67 तस्मादित्युत्तरस्य). Also used for limits and sources.
Indeclinable An avyaya such as च "and", वा "optionally", न "not".

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Full reading

नासिक-उदर-ओष्ठ-जङ्घा-दन्त-कर्ण-शृङ्गात् च

Pāṇini leaves out words that carry over from earlier sūtras. Expand to see the rule as a reader actually understands it.

Commentary

And the feminine ङीष् comes optionally after compounds ending with 1. नासिका 'a nose' 2. उदर 'belly' 3. ओष्ठ 'lip' 4. जंघा 'leg' 5. दन्त 'tooth' 6. कर्ण 'ear' and 7. शृङ्ग 'horn'.
This is an exception to the last and the succeeding sutra. Some of the words above given are either such which have a conjunct penultimate letter, and thus beyond the scope of 4.1.54; or they are polysyllabic words and thus covered by 4.1.56. Though this rule is an exception to 4.1.54 and 4.1.56; yet it is qualified by 4.1.57. Thus तुङ्गनासिका or तुङ्गनासिकी, तिलोदरी or तिलोदरा, बिम्बोष्ठी or बिम्बोष्ठा, दीर्घजंघी or दीर्घजंघा, समदन्ती or समदन्ता, चारुकर्णी or चारुकर्णा, तीक्ष्णसृङ्गी or तीक्ष्णशृङ्गा ॥

Vart:- So also after the word पुच्छ 'tail'. As कल्याणपुच्छा or कल्याणपुच्छी ॥

Vart:- The affix nish comes necessarily and not optionally after compounds of पुच्छ with कबर, मणि, विष and शर ॥ As, कबरपुच्छी 'a peahen' मणिपुच्छी, विषपुच्छी, 'a scorpion' and शरपुच्छी ॥

Vart:- So also the feminine affix nish is necessarily employed after the words पक्ष and पुच्छ when used as a simile or as objects of comparison. As उलूकपक्षी सेना, उलूकपुच्छी शाला ॥

Śrīśa Chandra Vasu, The Ashtadhyayi of Panini (1897), via ashtadhyayi.com.