त्रसिगृधिधृषिक्षिपेः क्नुः

Operational rule vidhi

The affix क्नु comes after the verbs - 1. त्रस् 'to tremble' 2. गृध् 'to be greedy' 3. धृष् 'to be bold' 4. क्षिप् 'to throw' in the sense of 'agent having such a habit etc.'.

Word by word (padaccheda)

Left context त्रसि- गृधि- धृषि- क्षिपेः pañcamī · ekavacanaSubstitute / term नु ः prathamā · ekavacana

Left context Pañcamī (5th case): "after X", the operation applies to what follows (1.1.67 तस्मादित्युत्तरस्य). Also used for limits and sources.
Substitute / term Prathamā (1st case): in operational rules, the substitute or the thing introduced; in definitions, the term or its referent.

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Pink letters are it-markers (anubandhas), tags dropped by 1.3.9. Hover or tap one for the rule that marks it, or try the it-letter finder.

Full reading

त्रसि-गृधि-धृषि-क्षिपेः क्नुः

Pāṇini leaves out words that carry over from earlier sūtras. Expand to see the rule as a reader actually understands it.

Commentary

The affix क्नु comes after the verbs - 1. त्रस् 'to tremble' 2. गृध् 'to be greedy' 3. धृष् 'to be bold' 4. क्षिप् 'to throw' in the sense of 'agent having such a habit etc.'.
As, त्रस्नुः 'trembling'; गृध्नुः 'greedy'; धृष्णुः 'bold'; and क्षिण्नुः 'throwing'.

Śrīśa Chandra Vasu, The Ashtadhyayi of Panini (1897), via ashtadhyayi.com.