जात्यन्ताच्छ बन्धुनि

Operational rule vidhi

The affix छ (ईय) is added without a change of sense to a stem ending in the word जाति when it means appropriateness.

Word by word (padaccheda)

Left context जात्यन्तात् pañcamī · ekavacanaWord छ Right context बन्धुनि saptamī · ekavacana

Left context Pañcamī (5th case): "after X", the operation applies to what follows (1.1.67 तस्मादित्युत्तरस्य). Also used for limits and sources.
Word Case not marked in the source data.
Right context Saptamī (7th case): "when X follows", the operation applies to what precedes (1.1.66 तस्मिन्निति निर्दिष्टे पूर्वस्य). Also a condition or domain ("in the sense of").

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Full reading

जात्यन्तात् छ बन्धुनि

Pāṇini leaves out words that carry over from earlier sūtras. Expand to see the rule as a reader actually understands it.

Commentary

The affix छ (ईय) is added without a change of sense to a stem ending in the word जाति when it means appropriateness.
The word बन्धु means the substance or substratum in which jati or generic quality adheres or any thing by which a thing is regulated and individualised (बध्यते नियम्यते स्वतन्त्री क्रियते). Thus बन्धु means that substance by which the genus Brahmana, Kshatriya &c is distinguished. Thus ब्राह्मणजातीयः "appropriate to a Brahmana". क्षत्रियजातीयः, वैश्यजातीयः &c.

Why do we say "when meaning appropriateness". Observe ब्राह्मणजातिः शोभना "the Brahmana caste is beautiful".

Śrīśa Chandra Vasu, The Ashtadhyayi of Panini (1897), via ashtadhyayi.com.