जिह्वामूलाङ्गुलेश्छः

Operational rule vidhi

After the words 1. जिह्वामूल and 2. अङ्गुलि there is the affix छ in the sense of 'who stays there'.

Word by word (padaccheda)

Left context जिह्वामूल- अङ्गुलेः pañcamī · ekavacanaSubstitute / term छः prathamā · ekavacana

Left context Pañcamī (5th case): "after X", the operation applies to what follows (1.1.67 तस्मादित्युत्तरस्य). Also used for limits and sources.
Substitute / term Prathamā (1st case): in operational rules, the substitute or the thing introduced; in definitions, the term or its referent.

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Full reading

जिह्वामूल-अङ्गुलेः छः

Pāṇini leaves out words that carry over from earlier sūtras. Expand to see the rule as a reader actually understands it.

Commentary

After the words 1. जिह्वामूल and 2. अङ्गुलि there is the affix छ in the sense of 'who stays there'.
This debars यत् 4.3.55. Thus जिह्वामूलीयम्, अंगुलीयम् ॥

Śrīśa Chandra Vasu, The Ashtadhyayi of Panini (1897), via ashtadhyayi.com.