मद्रेभ्योऽञ्

Operational rule vidhi

The affix अञ् is added to the word मद्र , preceded by a word denoting 'direction', in the remaining senses.

Word by word (padaccheda)

Left context मद्रेभ्यः pañcamī · bahuvacanaSubstitute / term अ prathamā · ekavacana

Left context Pañcamī (5th case): "after X", the operation applies to what follows (1.1.67 तस्मादित्युत्तरस्य). Also used for limits and sources.
Substitute / term Prathamā (1st case): in operational rules, the substitute or the thing introduced; in definitions, the term or its referent.

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Full reading

मद्रेभ्यः अञ्

Pāṇini leaves out words that carry over from earlier sūtras. Expand to see the rule as a reader actually understands it.

Commentary

The affix अञ् is added to the word मद्र , preceded by a word denoting 'direction', in the remaining senses.
Thus पौर्वमद्रः f. द्री; आपरमद्रः f. द्री ॥ Here contrary to the examples in the preceding sutra, the initial vowel is vriddhied by 7.3.13 which makes a special exception in the case of the word मद्र ॥

Śrīśa Chandra Vasu, The Ashtadhyayi of Panini (1897), via ashtadhyayi.com.