घञपोश्च

Operational rule vidhi

And when the affix घञ् and अप् follow घसॢ is the substitute of अद्.

Word by word (padaccheda)

Right context घञ्- अपोः saptamī · dvivacanaIndeclinable च avyaya

Right context Saptamī (7th case): "when X follows", the operation applies to what precedes (1.1.66 तस्मिन्निति निर्दिष्टे पूर्वस्य). Also a condition or domain ("in the sense of").
Indeclinable An avyaya such as च "and", वा "optionally", न "not".

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Full reading

घञ्-अपोः च

Pāṇini leaves out words that carry over from earlier sūtras. Expand to see the rule as a reader actually understands it.

Commentary

And when the affix घञ् and अप् follow घसॢ is the substitute of अद्।
घासः 'food, grass,' अद् + घञ् = घास + अ 3.3.19. प्रघसः 'voracity.' प्र + अद् + अप् = प्र + घस.

Śrīśa Chandra Vasu, The Ashtadhyayi of Panini (1897), via ashtadhyayi.com.