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स्थानेऽन्तरतमः The nearest substitute

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Which guṇa?

A rule says:

इ → guṇa

guṇa = अ, ए, ओ (1.1.2 अदेङ् गुणः). Which one replaces इ?

The answer comes from 1.1.50 स्थानेऽन्तरतमः: "in a substitution, the nearest one". Rules such as 7.3.84 just say "guṇa"; 1.1.50 picks the member.

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